Paul is the Stephen A. Levin Family Dean of the College of Arts and Sciences and a Professor of Biology at the University of Pennsylvania, where he has been a faculty member since 1997. He is a member of the graduate groups in Biology, Genomics and Computational Biology, and History and Philosophy of Science. He earned his undergraduate degree at the Indiana University School of Music, where he received a B.Mus. in Violin Performance. He obtained his PhD in Evolutionary Biology in 1993 at the University of Chicago under Brian Charlesworth and did postdoctoral work at Michigan State University with Richard Lenski, where he conducted studies on the evolution of genetic mutation rates that have continued here at Penn. His honors include a Sloan Foundation Outstanding Young Investigator award for studies of molecular evolution, the Keeley Visiting Fellowship at Wadham College, Oxford, and the Ira H. Abrams Award for teaching in the School of Arts and Sciences at the University of Pennsylvania.